Video clip
Curators at the Scottish National Gallery provide insights into Titian's "Diana and Callisto" and "Diana and Actaeon", and the Venetian artist's pioneering work in the use of colour oils.
Titian's large painting depicting Actaeon stumbling on Diana bathing reveals his "sumptuous, senuous", broad-brush style that was very different from his Northern European contemporaries.
Permanent exhibitions at Edinburgh galleries are free to view.
National Gallery of Scotland and National Gallery in London purchase Diana and Callisto.